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5		-= **The cortical mesocircuit model** =
6		-
7		-A layered network model covering 4x4 mm² of cortical surface area
	5	+= A mesoscopic layered cortical network model =
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	13	+= [[image:mesocircuit_sketch.png]] =
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17		-= Overview =
	15	+= What can I find here? =
18	18
19		-The cortical mesocircuit model is a lateral extension of the microcircuit model published by [[Potjans & Diesmann (2014)>>https://doi.org/10.1093/cercor/bhs358]].
	17	+* Notice how the table of contents on the right
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22		-[[image:mesocircuit_sketch.png||height="333" width="500"]]
	21	+= Who has access? =
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	23	+Describe the audience of this collab.
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27	27	{{box title="**Contents**"}}
28	28	{{toc/}}
29	29	{{/box}}
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34		-Like the microcircuit, the mesocircuit represents four layers of cortex, L2/3, L4, L5, and L6, each consisting of a population of excitatory neurons and a population of inhibitory neurons. External input can be provided by a thalamic population TC.
35		-
36		-The mesocircuit is a spatially structured network model covering 4x4 mm² of cortical surface area which is similar in size to multi-electrode arrays in use today.
37		-
38		-The neurons of each population are assigned random positions in two-dimensional layers and they are connected with distance-dependent connection probabilities and conduction delays.
39		-
40		-The densities of neurons and local synapses in the network are assumed to be realistic, amounting to ~~1.2 million neurons and ~~5.5 billion synapses.
41		-
42		-The model accounts for both spiking activity simulated with [[NEST>>https://www.nest-simulator.org]] and local-field potentials relying on [[LFPy>>https://LFPy.rtfd.io]].
43		-
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45		-= Status of model development =
46		-
47		-The mesocircuit is still under development and the neither the code nor a peer-reviewed manuscript have been published, yet.
48		-
49		-To date, only a [[preprint>>https://arxiv.org/abs/1805.10235]] with detailed model documentation and preliminary results is available on arXiv.org.
50		-
51		-The Drive of this Collab provides access to a [[recent version of the model implementation>>https://wiki.ebrains.eu/bin/view/Collabs/mesocircuit/Drive]].
52		-
53		-The development of the mesocircuit model belongs to task "SGA2 T4.2.1 - Spiking mesoscale cortical models with spatial organization" and constitutes the [[component C2418>>https://plus.humanbrainproject.eu/components/2418]].
54		-
55		-The main contributors are Johanna Senk, Espen Hagen, Sacha J. van Albada and Markus Diesmann.
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